Chrysalis Snails s.l. Pupilloidea Turton, 1831

Diese Seite auf Deutsch!

In the superfamily of Chrysalis Snails s.l., there is a number of related snail familiess, some of which are described on the present homepage.

Mountain Bulin Snail (Ena montana - Enidae). Photo: © Stefan Haller, (schneckenfoto.ch).		Large Barrel Snail (Orcula dolium - Orculidae). Photo: Olivier Gargominy (Source).
Moss Chrysalis Snail (Pupilla muscorum - Pupillidae). Photo: © Stefan Haller (schneckenfoto.ch).		Ribbed Grass Snail (Vallonia costata - Valloniidae). Photo: © Stefan Haller (schneckenfoto.ch).

According to MolluscaBase (see below), presently those are (Note: Families not (yet) described on the present homepage are greyed out):

• Family Achatinellidae Gulick, 1873
• Family Agardhiellidae Harl & Páll-Gergely, 2017
• Family Amastridae Pilsbry, 1910
• Family Argnidae Hudec, 1965  ().
• Family Cerastidae Wenz, 1923
• Family Cochlicopidae Pilsbry, 1900 (1879) - Pillar Snails.
• Family Draparnaudiidae Solem, 1962
• Family Enidae B. B. Woodward, 1903 (1880) - Bulin Snails.
• Family Fauxulidae Harl & Páll-Gergely, 2017
• Family Gastrocoptidae Pilsbry, 1918
• Family Hypselostomatidae Zilch, 1959
• Family Lauriidae Steenberg, 1925 - Chrysalis Snails (1).
• Family Odontocycladidae Hausdorf, 1996
• Family Orculidae Pilsbry, 1918 ().
• Family Pagodulinidae Pilsbry, 1924  ().
• Family Partulidae Pilsbry, 1900 - Pacific Tree Snails
• Family Pleurodiscidae Wenz, 1923
• Family Pupillidae W. Turton, 1831 - Chrysalis Snails s.str. (2).
• Family Pyramidulidae Kennard & B. B. Woodward, 1914  ().
• Family Spelaeoconchidae A. J. Wagner, 1928
• Family Spelaeodiscidae Steenberg, 1925
• Family Strobilopsidae Wenz, 1915
• Family Valloniidae E. S. Morse, 1864 - Grass Snails.
• Family Vertiginidae Fitzinger, 1833 - Whorl Snails.

MolluscaBase eds. (2021): Pupilloidea W. Turton, 1831.

However, the Orculidae have undergone a major revision in recent time. The paper by Harl et al. (2017), mentioned below, on the basis of threee mitochondrial and two nuclear genes, came to the conclusion that several genera previously listed among the Orculidae, in fact constitute separate groups. So today only the genera Alvariella, Orcula, Orculella, Pilorcula, Schileykula and Sphyradium are counted among the Orculidae. Two new families, Odontocycladidae and Pagodulinidae, were formed outside of the Orculidae family. Besides, the family Fauxulidae is freshly established.

• Harl, J., Haring, E., Asami, T., Sittenthaler, M., Sattmann, H., Páll-Gergely, B. (2017): Molecular systematics of the land snail family Orculidae reveal paraphyly and deep splits within the clade Orthurethra (Gastropoda: Pulmonata). Zoological Journal of the Linnean Society. 181(4): pp. 778-794. (Link).
• Bouchet, P.; Rocroi, J.P.; Hausdorf, B.; Kaim, A.; Kano, Y.; Nützel, A.; Parkhaev, P.; Schrödl, M; Strong, E.E. (2017): Revised classification, nomenclator and typification of gastropod and monoplacophoran Families. Malacologia. 61(1-2): 1-526.

Additionally the family Chondrinidae, together with the Truncatellinidae, is placed in a separate superfamily, the Chondrinoidea, outside of the Pupilloidea.

• MolluscaBase eds. (2025): Chondrinoidea Steenberg, 1925.
• Saadi, A. J.; Mordan, P. B.; Wade, C. M. (2021): Molecular phylogeny of the Orthurethra (Panpulmonata: Stylommatophora). Zoological Journal of the Linnean Society. (Abstract). 
  

With photos by Stefan Haller: http://www.schneckenfoto.ch.

Latest Change: 25.10.2026 (Robert Nordsieck).